Geomitridae is a taxonomic family of small to medium-sized air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Helicoidea.
Anatomy
The family is characterized by the presence of a free right ommatophoral retractor (passing outside the peni-oviducal angle) and a double stimulatory apparatus. A free right ommatophoral retractor has been linked to adaptation to xeric habitats
Taxonomy
The family Geomitridae is subdivided in the following subfamilies (according to Razkin et al., 2015) Subfamily Geomitrinae tribe Cochlicellini Schileyko, 1972 tribe Geomitrini C. Boettger, 1909 - synonym: Ochthephilinae Zilch, 1960 (n.a.) tribe Ponentinini Razkin et al., 2015 Subfamily Helicellinae tribe Cernuellini Schileyko, 1991 tribe Helicellini Ihering, 1909 - synonym: Jacostidae Pilsbry, 1948 (inv.) tribe Helicopsini H. Nordsieck, 1987 tribe Plentuisini Razkin et al., 2015 tribe Trochoideini H. Nordsieck, 1987
Genera
Genera with the family Geomitridae include:
Not belonging to a subfamily: Keraea Gude, 1911 Peridotitea Torres Alba, Holyoak, D. T., Holyoak, G. A., Vázquez Toro & Ripoll, 2018
Subfamily Geomitrinae
tribe Cochlicellini
Cochlicella Férussac, 1821
Monilearia Mousson, 1872
Obelus W. Hartmann, 1842
Ripkeniella Hutterer & E. Gittenberger, 1998
tribe Geomitrini
Actinella R. T. Lowe, 1852
Callina R. T. Lowe, 1855
Caseolus R. T. Lowe, 1852
Discula R. T. Lowe, 1852
Disculella Pilsbry, 1895
Domunculifex Brozzo, De Mattia, Harl & Neiber, 2020
Geomitra Swainson, 1840
Helicomela R. T. Lowe, 1855
Heterostoma W. Hartmann, 1843: synonym of Steenbergia Mandahl-Barth, 1950 (invalid: junior homonym of Heterostoma de Filippi, 1837 [Platyhelminthes]) Hystricella R. T. Lowe, 1855 Lemniscia R. T. Lowe, 1855 Moreletina de Frias Martins, 2002 Plebecula R. T. Lowe, 1852 Pseudocampylaea L. Pfeiffer, 1877 Serratorotula Groh & Hemmen, 1986 Spirorbula R. T. Lowe, 1852 Steenbergia Mandahl-Barth, 1950 Testudodiscula Brozzo, De Mattia, Harl & Neiber, 2020 Wollastonaria De Mattia, Neiber & Groh, 2018 tribe Ponentinini Ponentina P. Hesse, 1921
subfamily Helicellinae
tribe Cernuellini
Alteniella Clerx & Gittenberger, 1977
Cernuella Schlüter, 1838
Microxeromagna Ortiz de Zárate López, 1950
Xeroplana Monterosato, 1892
Xerosecta Monterosato, 1892
tribe Helicellini
Backeljaia Chueca et al., 2018
Candidula Kobelt, 1871
Helicella Férussac, 1821
Orexana Chueca et al., 2018
Xerogyra Monterosato, 1892
Xeroleuca Kobelt, 1877
Xeroplexa Monterosato, 1892
Xerotricha Monterosato, 1892
Zarateana Chueca et al., 2018
tribe Helicopsini H. Nordsieck, 1987
Helicopsis Fitzinger, 1833
Pseudoxerophila Westerlund, 1879
Xerolenta Monterosato, 1892
Xeromunda Monterosato, 1892
Xeropicta Monterosato, 1892
tribe Plentuisini
Plentuisa Puente & Prieto, 1992
tribe Trochoideini
Trochoidea T. Brown, 1827
Xerocrassa Monterosato, 1892
References
Bouchet P., Rocroi J.P., Hausdorf B., Kaim A., Kano Y., Nützel A., Parkhaev P., Schrödl M. & Strong E.E. (2017). Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia. 61(1-2): 1-526
External links
https://liferecovernatura.madeira.gov.pt
Razkin, O., Gómez-Moliner, B. J., Prieto, C. E., Martínez-Ortí, A., Arrébola, J. R., Muñoz, B., Chueca, L. J. & Madeira, M. J. (2015). Molecular phylogeny of the western Palaearctic Helicoidea (Gastropoda, Stylommatophora). Molecular Phylogenetics and Evolution. 83: 99-117