If the fact that males fight before they are paired and in the absence of a female could be placed beyond all question, it would no longer be possible to regard her possession as the end for which they are contending, and consequently there would be no need to produce further evidence. But the examples which I have given refer, of course, to only a few migrants and a few residents--and moreover it must be admitted that a female _is_ often conspicuous during the battles--so that by themselves they must be regarded, and rightly so, as inconclusive. We must therefore pass on to consider evidence of a somewhat different character.
I spoke of the complexity of the strife. By this I mean that it is not merely a matter of disputes between adjoining males, but that it is a far more comprehensive business involving both sexes. Thus female fights with female and pair with pair, or a male will attack a female, or, again, a pair will combine against a single male or a single female. And from all this complexity of strife we gain much valuable evidence in regard to the question immediately before us. For when one pair attacks another, or males that are definitely paired fight with one another, or an unpaired male attacks either sex of a neighbouring pair indiscriminately, there is surely little ground for supposing that the possession of a mate is the reason of it all.
The battles between pairs of the same species are by no means uncommon. Observe, for example, the central pair of three pairs of Reed-Buntings occupying adjoining territories, and keep a daily record of the routine of activity practised by both sexes during the early hours of the morning; then, at the close of the season, summarise all the fighting under different headings, and it will be found that the number of occasions upon which the central pair attacked, or was attacked by, neighbouring pairs will form a considerable portion of the whole.
Or watch the Moor-Hen, and for the purpose choose some sheet of water large enough to accommodate three or more pairs, and so situated that the birds can always be kept in view. Early in February the pool will be haunted by numbers of individuals of both sexes, all swimming about together, and, if the pool is surrounded by arable land, wandering over that land subject to no territorial restrictions, apparently free to seek food where they will. But as time goes by, their number gradually decreases until a few pairs only remain, and these will occupy definite areas. If careful watch is then kept and the relations of the pairs closely studied, there will be no difficulty in observing the particular kind of warfare to which I am alluding, and it will be noticed that the encounters are of a particularly violent description. Thus two pairs approach one another, and, when they meet, throw themselves upon their backs, each bird striking at its adversary with its feet or seizing hold of it with its beak; and though, in the commotion that ensues, it is almost impossible to determine what exactly is happening, there is reason to believe that the sexes attack one another indiscriminately.