As a proof of leaf variegation being a disease, the speaker alluded to cited a case in which a green leaved abutilon, upon which a variegated leaved variety had been grafted, threw out a variegated leaved shoot below the graft. This can easily be explained. The growth of the trunk or stem of all exogenous plants, or those which increase in size on the outside of the stem, is brought about by the descent of certain formative tissue called cambium, elaborated by the leaves and descending between the old wood and the bark, where it is formed into alburnum or woody matter. Some think that it is also formed by the roots and ascends from them as well as descending from the leaves. Be this as it may, there is no doubt about its descent. In such comparatively soft-wooded, free growing plants as the abutilon the descent of the cambium is very free and in considerable quantity, so that the stock would soon be inclosed in a layer of it descending from the graft. When being converted into woody matter it also forms adventitious buds which under certain favorable circumstances will emit shoots of the same character as the graft from which it was derived. The graft is such cases may be said to inclose the stock in a tube of its own substance, leaving the stock unaffected otherwise. The variegated shoot in this case was in reality derived from the downward growth of the graft and not from the original stock, which was not therefore contaminated by the graft. In cases where the stock is of much slower growth than the graft, or the graft is inserted upon a stock of some other species, the descending cambium does not inclose the stock, but makes layers of wood on the stem of the graft, which thus, as is frequently seen, overgrows the stock, sometimes to such an extent as to make it unsightly. Nobody ever saw an apple shoot from a crab stock, a pear from a quince stock, or a peach shoot from a plum stock. This is one of the arguments in favor of the view that cambium also rises from the roots.
Again, to show that the stock is not affected by the graft, or the graft by the stock, except as to root power, let any person graft a white beet upon a red beet, or contrariwise, when about the size of a goosequill, and when they have attained their full growth, by dividing the beet lengthwise he will find the line of demarkation between the colors perfectly distinct, neither of them running into the other.