The most celebrated companions of Polygnotos, but, as Ælian remarks, not equalling him in greatness, were Micon of Athens, whose name has already been mentioned, and Panainos, a cousin of Pheidias, who, besides the battle of Marathon in the Poikile, executed the paintings upon the throne of the Pheidian Zeus in Olympia. Dionysios of Colophon and Pauson have already been spoken of. The first seems to have carried out the strict carefulness of his model, Polygnotos, to a degree which was naturally unfavorable alike to grace and to greatness of style. Pauson, though accounted an artist by Aristotle, may be compared to Buffalmacco, scorned and derided, among the companions of Giotto; not fitted for productions of a grand style, he did not attempt them, and his nude paintings, without ethical significance, were harmful to young observers.
Among the other distinguished masters of this time, Calliphon appears most nearly to have followed in the footsteps of Polygnotos; but his brother Aristophon, who brought painting upon panels into general use, pursued technical methods opposed to this school. The style of Polygnotos was also abandoned by the Samian Agatharchos, a self-instructed decorator and scene-painter who, in an essay upon scenographic painting, established principles upon which, after his time, this art was further developed. In scene-painting the indispensable aim after illusory appearances must have led to the observation and imitation of the effect of more or less light--that is to say, of paler or deeper shades in the local color--and thus have brought painting to a point of development not hitherto attained by any nation of antiquity.
The important advance indicated by Agatharchos in scenography was made in the painting of figures by Apollodoros of Athens. The accounts of him are few, and in part incomprehensible; but Plutarch says plainly that he discovered the mixing of colors and the variation of shade upon them, and Pliny calls him the first master of illusion. Strictly speaking, he was not the sole author of the innovation, since Agatharchos went before him; and if he received the cognomen of _skiagraphos_--painter in light and shade--it must be understood that the word skiagraphia was used to signify scenography. But he was, at all events, the first to apply these principles to figure-painting, developing a treatment quite different from that employed in the architectural painting so extensively in use for the stage. The important result of this innovation may well be imagined, and it is not strange that the ground thus gained should have been promptly occupied by other masters of the art, who rapidly brought painting to a perfection almost equal to that of sculpture.