The Sahara pump theory is a hypothesis that explains how flora and fauna migrated between Eurasia and Africa via a land bridge in the Levant region. It posits that extended periods of abundant rainfall lasting many thousands of years (pluvial periods) in Africa are associated with a "wet-green Sahara" phase, during which larger lakes and more rivers existed. This caused changes in the flora and fauna found in the area. Migration along the river corridor was halted when, during a desert phase 1.8–0.8 million years ago (mya), the Nile ceased to flow completely and possibly flowed only temporarily in other periods due to the geologic uplift (Nubian Swell) of the Nile River region.
Mechanism
During periods of a wet or Green Sahara, the Sahara and Arabia become a savanna grassland and African flora and fauna become common. Following inter-pluvial arid periods, the Sahara area then reverts to desert conditions, usually as a result of the retreat of the West African Monsoon southwards. Evaporation exceeds precipitation, the level of water in lakes like Lake Chad falls, and rivers become dry wadis. Flora and fauna previously widespread as a result retreat northwards to the Atlas Mountains, southwards into West Africa, or eastwards into the Nile Valley and thence either southeast to the Ethiopian Highlands and Kenya or northeast across the Sinai into Asia. This separates populations of some of the species in areas with different climates, forcing them to adapt, possibly giving rise to allopatric speciation.
Plio-Pleistocene
The Plio-Pleistocene migrations to Africa included the Caprinae in two waves at 3.2 Ma and 2.7–2.5 Ma; Nyctereutes at 2.5 Ma, and Equus at 2.3 Ma. Hippotragus migrated at 2.6 Ma from Africa to the Siwaliks of the Himalayas. Asian bovids moved to Europe and to and from Africa. The primate Theropithecus experienced contraction and its fossils are found only in Europe and Asia, while Homo and Macaca settled wide ranges.
185,000–20,000 years ago
Between about 133 and 122 thousand years ago (kya), the southern parts of the Saharan-Arabian Desert experienced the start of the Abbassia Pluvial, a wet period with increased monsoonal precipitation, around 100-200 mm/year. This allowed Eurasian biota to travel to Africa and vice versa. The growth of speleothems (which requires rainwater) was detected in Hol-Zakh, Ashalim, Even-Sid, Ma'ale-ha-Meyshar, Ktora Cracks, Nagev Tzavoa Cave. In Qafzeh and Es Skuhl caves, where at that time precipitation was 600–1000 mm/year, the remains of Qafzeh-Skhul type anatomically modern humans are dated from this period, but human occupation seems to end in the later arid period.
The Red Sea coastal route was extremely arid before 140 and after 115 kya. Slightly wetter conditions appear at 90–87 kya, but it still was just one tenth the rainfall around 125 kya. Speleothems are detected only in Even-Sid-2.
In the southern Negev Desert speleothems did not grow between 185–140 kya (MIS 6), 110–90 (MIS 5.4–5.2), nor after 85 kya nor during most of the interglacial period (MIS 5.1), the glacial period and Holocene. This suggests that the southern Negev was arid to hyper-arid in these periods.
The coastal route around the western Mediterranean may have been open at times during the last glacial; speleothems grew in Hol-Zakh and in Nagev Tzavoa Caves. Comparison of speleothem formation with calcite horizons suggests that the wet periods were limited to only tens or hundreds of years.
From 60–30 kya there were extremely dry conditions in many parts of Africa.
Last Glacial Maximum
An example of the Saharan pump has occurred after the Last Glacial Maximum (LGM). During the Last Glacial Maximum the Sahara desert was more extensive than it is now with the extent of the tropical forests being greatly reduced. During this period, the lower temperatures reduced the strength of the Hadley Cell whereby rising tropical air of the Inter-Tropical Convergence Zone (ITCZ) brings rain to the tropics, while dry descending air, at about 20 degrees north, flows back to the equator and brings desert conditions to this region. This phase is associated with high rates of wind-blown mineral dust, found in marine cores that come from the north tropical Atlantic.