In western British Columbia, Washington, and Oregon no evidences of intergradation between _S. v. vagrans_ and the races _setosus_, _permiliensis_, _bairdi_, _yaquinae_, or _pacificus_ are seen. In this region _S. v. vagrans_ occurs sympatrically with one or the other of these subspecies. Different degrees of differentiation thus obtain between the subspecies _vagrans_ as here defined and the surrounding subspecies of _Sorex vagrans_ to wit: complete intergradation and allopatry in Utah with _S. v. obscurus_; partial intergradation and partial sympatry with _S. v. obscurus_ in the foothill region from Idaho to British Columbia; no intergradation and complete sympatry with all the other races of _Sorex vagrans_ from the Cascades to the coast and south to San Francisco Bay. The relationship of _S. v. vagrans_ to the wandering shrews of the high Sierra is discussed on page 58.

Throughout most of the Great Basin and Columbian Plateau _Sorex vagrans_ is, with the exception of the rare _S. merriami_ and _S. preblei_, the only small shrew. In the Cascades and in the coastal lowlands it is the only small shrew except for _S. cinereus_ and _S. trigonirostris_, both extremely rare and local in this region. _S. vagrans_ seemingly competes to a certain extent with the larger _S. trowbridgii_ in western Washington and seems to be partially dominant to _trowbridgii_, at least in marshy habitats (Dalquest, 1941:171).

Speciation of the Wandering Shrew — Part 46

_Specimens examined._--Total number, 1197.

Speciation of the Wandering Shrew — Part 46

BRITISH COLUMBIA: _Osoyoos District_: Okanagan, 20 PMBC; Okanagan Landing, 2 PMBC; Nahun Plateau, 2 PMBC. _Vancouver District_: Vancouver, 2 PMBC. _New Westminister District_: Port Moody, 16 BS; Westminster Jct., 4 AMNH; Langley, 1 BS; Vedder Crossing, 1 PMBC; Huntingdon, 69 NMC; Sumas, 16 BS; Cultus Lake, 1 NMC. _Similkameen District_: Princeton, 6 Mile Creek, 1 NMC. Hedley, Stirling Creek, 7 NMC; Fairview-Keremeos Summit, 5 NMC; Oliver, 1 NMC; Westbridge, 6 NMC; Osoyoos, 1 PMBC; Osoyoos-Bridesville Summit, 4 NMC; Cascade, 7 NMC. _Nelson District_: Kuskonook, 1 PMBC; Rossland, 14 NMC; Trail, 2 NMC; Creston, 4 PMBC, 4 NMC; near Creston, 7 NMC. _Cranbrook District_: Cranbrook, 5 BS; Yahk, 2 NMC; Yahk Camp 6, 2 NMC; Goatfell, 2 NMC. _Fernie District_: Newgate, 3 NMC.

Speciation of the Wandering Shrew — Part 46