_Habit_ has a decided influence towards inducing variation. As the blacksmith's right arm becomes more muscular from the habit of exercise induced by his vocation, so we find in domestic animals that use, or the demand created by habit, is met by a development or change in the organization adapted to the requirement. For instance, with cows in a state of nature or where required only to suckle their young, the supply of milk is barely fitted to the requirement. If more is desired, and if the milk be drawn completely and regularly, the yield is increased and continued longer. By keeping up the demand there is induced in the next generation a greater development of the secreting organs, and more milk is given. By continuing the practice, by furnishing the needful conditions of suitable food, &c., and by selecting in each generation those animals showing the greatest tendency towards milk, a breed specially adapted for the dairy may be established. It is just by this mode that the Ayrshires have, in the past eighty or a hundred years, been brought to be what they are, a breed giving more good milk upon a given quantity of food than any other.

It is because the English breeders of modern Short-horns altogether prefer beef-making to milk-giving properties that they have constantly fostered variation in favor of the one at the expense of the other until the milking quality in many families is nearly bred out. It was not so formerly--thirty years ago the Short-horns (or as they were then usually called, the Durhams) were not deficient in dairy qualities, and some families were famous for large yield. By properly directed efforts they might, doubtless, be bred back to milk, but of this there is no probability, at least in England, for the tendency of modern practice is very strong toward having each breed specially fitted to its use--the dairy breeds for milk and the beef breeds for meat only. The requirements of the English breeder are in some respects quite unlike those of New England farmers--for instance, as they employ no oxen for labor there is no inducement to cultivate working qualities even, in connection with beef.

The Principles of Breeding — Part 18

As an illustration of the effect of habit, Darwin[5] cites the domestic duck, of which he says, "I find that the bones of the wing weigh less, and the bones of the leg more, in proportion to the whole skeleton, than do the same bones in the wild duck; and I presume that this change may be safely attributed to the domestic duck flying much less and walking more than its wild parent." And again, "not a single domestic animal can be named which has not in some country drooping ears, and the view suggested by some authors, that the drooping is due to the disuse of the muscles of the ear, from the animals not being much alarmed by danger, seems probable."

The Principles of Breeding — Part 18