The general shape of the palatoquadrate complex is most nearly like that of _Rhabdoderma elegans_ (Moy-Thomas, 1937:fig. 5). The orientation of the complex in the living fish was probably oblique, with the processus ascendens nearly vertical, the quadrate oblique, and the ventral process of the endopterygoid extending dorsoanteriorly and articulating with the parasphenoid.
Of the hyoid arch only the ceratohyals (see fig. 5A) are preserved in several specimens. These are long, curved bones with a posteroventral process and widened, flaring posterior margin. The medial (?) surface is concave in one specimen. The lateral (?) surface displays a distinct ridge on several specimens, arising on the dorsal surface opposite the posteroventral process and extending diagonally to the anteroventral end of the anterior limb. The impression of one other specimen appears to have a central ridge because of greater dorsal thickness and narrowness. Both surfaces are unornamented.
The urohyal (see fig. 5B) is an unornamented, Y-shaped bone, with the stem of the Y pointing anteriorly. Orientation with respect to dorsal and ventral surfaces is uncertain. In one view a faint ridge, also Y-shaped, occurs on the expanded posterior portion, and the surface is convex. The anterior process has a convex surface, sloping evenly off to the lateral margin; the opposite side of the process has a concave surface. The posterior portion has a slightly depressed area (see fig. 5B) at the junction of the "arms" of the Y.
The five branchial arches are represented by the ceratobranchials, several of which are preserved on K. U. no. 11431. These are long bones with anteriorly curving ventral ends. The medial surfaces are partly covered with minute granular teeth; only the dorsal part is without teeth. The dorsal articular surface is convex dorsally and rounded.
The mandible (fig. 3), the best specimens of which are K. U. nos. 788 and 11425, is seen only in lateral and ventral views, with only angular, splenial and dentary visible.
The angular forms the main body of the mandible, and is similar to that of _Spermatodus_. The dorsal margin of the angular is expanded in the central region, with some variation. One specimen has an expanded portion slightly anterior to that of the opposite angular. The articular surface near the posterior end has not been observed; the posterior end of the angular slopes off abruptly. The anterior sutures are seen in only two specimens, K. U. nos. 788, 11425. The dentary meets the angular in a long oblique suture; the dentary gradually tapers posterodorsally and ends on the dorsal surface of the angular. The splenial fits into a posteriorly directed, deep V-shaped notch on the ventral surface. The lateroventral surface of the angular contains sensory pores of the mandibular line. The ventral surface extends medially into a narrow shelf, approximately 1.0 mm. wide, which extends the full length of the bone; the external surface of this shelf is smooth and slightly concave dorsally. Ornamentation of the angular consists of tubercles and longitudinal or oblique striae, occurring mostly on the expanded portion. The medial surface is not seen. Several broken specimens show a central canal filled with a rod of calcite; in one of these the sensory pores are also calcite-filled and appear to be connected to the rod. Thus the pores originally opened into a central canal.