In _Delphinium_, as shown by Braun,[376] the stamens and carpels are members of a continuous spiral series, and in the double balsam an extra corolline whorl is produced, without the suppression of the stamens, in the following manner: the ordinary stamens are replaced by petals, the carpels by stamens, while an additional whorl of carpels is produced at the summit of the axis. In this instance, therefore, the doubling is distinctly referrible to an absolute increase in the number of whorls, and not to chorisis.[377]

On the other hand, it must be admitted that there are many cases which are not to be explained in any other way than that suggested by the French botanists before alluded to. Probably, the main difficulty in the way of accepting the doctrine of chorisis is the unfortunate selection of the word used to designate the process; this naturally suggests a splitting of an organ already perfectly formed into two or more portions, either in the same plane as the original organs, "parallel chorisis;" or at right angles to it "collateral chorisis." Indeed, before so much attention had been paid to the way in which the floral organs are developed, it was thought that an actual splitting and dilamination did really take place; Dunal and Moquin both assert as much. The truth would rather seem to be that, in the so-called parallel chorisis at least, the process is one of hypertrophy and over-development rather than of splitting. The adventitious petal or scale is an excrescence or an outgrowth from the primary organ, and formed subsequently to it.

Vegetable Teratology — Part 219

In the case of "compound stamens" the original stamens are first developed each from its own cellular "mamelon," or growing point; and, after a time, other secondary growing points emerge from the primary one, and in this way the stamens are increased in number, without reference, necessarily, to the so-called law of alternation. Outgrowths from leaves, multiplying the laminar surface, are alluded to under the head of hypertrophy, and it is probable that some of the cases of duplication of the flower, or of the formation of adventitious segments outside the ordinary corolla as alluded to in succeeding paragraphs (see Pleiotaxy of the corolla), are due to a similar process.[378]

Vegetable Teratology — Part 219