Dryopidae is a family of beetles, commonly named long-toed water beetles, in the superfamily Byrrhoidea. It was described by Gustaf Johan Billberg in 1820.

Description

Long-toed water beetles are named for their extended claws. Adults have dense hairs, which allow the beetles to breathe while underwater. The flight muscles of the females weaken as they age.

Dryopidae

When the pupae complete the imago stage of their life cycle, they move towards running water, and may be attracted to lights.

Dryopidae

Despite being referred to as aquatic insects, the beetles are unable to swim, clinging to detritus that float. All long-toed water beetles feed on plants that are in the water, but the larvae generally are terrestrial, and at least some also feed on plant matter. Stygoparnus is the only genus in the family in which both the larvae and adults are aquatic.

Dryopidae

Genera of the family closely resemble riffle beetles, but the antennae are different from the long-toed water beetles, looking similar to clubs.

Habitat

Members of this family are found on every continent, except Antarctica and Australia, being more common in the tropics. In 2005, a possible fossilized specimen of Dryopidae was found in the Crato Formation by entomologists David Grimaldi and Michael S. Engel.

Beetles that have water as a habitat, including long-toed water beetles, can help show the quality of fresh water.

Genera

Ahaggaria Bollow, 1938

Ceradryops Hinton, 1937

Drylichus Heller, 1916

Dryops Olivier, 1791

Elmomorphus Sharp, 1888

Elmoparnus Sharp, 1882

Geoparnus Besuchet, 1978

Guaranius Spangler, 1991

Helichus Erichson, 1847

Holcodryops Spangler, 1987

Malaiseianus Bollow, 1940

Onopelmus Spangler, 1980

Oreoparvus Delève, 1965

Pelonomus Erichson, 1847

Phallodryops Delève, 1963

Postelichus Nelson, 1989

Protoparnus Sharp, 1883

Pseudopelonomus Brown, 1981

Quadryops Perkins & Spangler, 1985

Rapnus Grouvelle, 1899

Sostea Pascoe, 1860

Sosteamorphus Hinton, 1936

Strina Redtenbacher, 1867

Stygoparnus Barr & Spangler, 1992

Uenodryops Satô, 1981

Palaeoriohelmis Bollow, 1940

Potamophilites Haupt, 1956

References

Byrrhoidea

Beetle families