98. _The Exoplasm of the Actipylea._--The extracapsular protoplasm of the ACANTHARIA or ACTIPYLEA differs in several important respects from that of other { lxvi}Radiolaria, and appears to undergo more significant differentiations than that of the three other legions. Since the pores in the wall of the central capsule are not distributed evenly and at equal intervals over its whole surface (as in the PERIPYLEA), but rather exhibit a regular disposition in groups at unequal intervals, the number of projecting pseudopodia is much less and the law of their arrangement different from that which obtains in the PERIPYLEA (§ 58). In many and probably in all ACANTHARIA they are divided into two groups, those which arise from the centre of the capsule and possess firm axial threads, and those which have not these characters (compare § 95, A). The axopodia, or stiff pseudopodia with axial threads, arise from the centre of the capsule, are present in much smaller numbers than the soft and flexible myxopodia, and are regularly disposed between the radial bars of acanthin, usually so that they are as far removed from them as possible, _i.e._, in the centre between each three or four bars; these latter may indeed be regarded as strongly developed axial threads, which have become changed into acanthin (§ 95, A). The soft myxopodia, or pseudopodia without axial threads, are much more numerous than the others, and arise from the sarcodictyum or exoplasmic network which ramifies over the surface of the calymma. Their number and arrangement seem, however, in many (if not in all) ACANTHARIA to be regular and not to possess the extraordinary variability seen in the other three legions. In many #Acanthometra# the sarcodictyum exhibits a symmetrical conformation, with regular or subregular, polygonal (mostly hexagonal) meshes, and generally the stronger threads of the sarcodictyum secrete a firm, homogeneous or fibrillar, striated substance, which forms a network of ridges on the surface of the calymma. In the #Acanthophracta# the place of this is taken by the acanthin network of the primary lattice-shell. The axopodia of the #Acanthometra# are usually about as long as the radial spines between which they stand; their stiff axial thread is surrounded by a soft sheath of protoplasm, communicating with the thin sarcomatrix which surrounds the central capsule. Numerous branches pass into the calymma from the exoplasmic sheath of the axial threads, and form by their interweaving a loose sarcoplegma. The most peculiar differentiated products of the exoplasm of the ACANTHARIA, however, are the myophane fibrillæ of the #Acanthometra#, which have already been described under the name of myophriscs (§ 96).